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Grammatocaulis donponii new genus new species a permineralized fern from the jurassic of queensland australia


, : Grammatocaulis donponii new genus new species a permineralized fern from the jurassic of queensland australia. Review of Palaeobotany & Palynology 66(1-2): 147-158

A permineralized fern stem from Middle to Late Jurassic sediments (Injune Creek Group) in the Wandoan area of southeastern Queensland is assigned to the new genus and species GRAMMATOCAULIS donponii. The stem possesses an ectophloic siphonostele with the xylem divided into an outer zone of long tracheids and an inner zone of short, wide, parenchyma-like tracheids surrounding a pith. Leaf traces are round in cross section in the parenchymatous inner cortex, becoming linear to oblong in the outer cortex and outside of the stem. Petiole bases have sclerotic rings around vascular strands, but lack stipular wings. Some of its outermost petiolar vascular strands assume a C-shape which may be due to distortion. Parenchymatous tissue occurs in the middle of the protoxylem at the ends of the elliptical petiolar vascular strands. At higher levels, the parenchymatous tissue of the strands is found in the metaxylem as well. Nonglandular multicellular scales occur on the petioles. Although the family of this genus is unknown, it is similar anatomically to Grammatopteris of the Anachoropteridaceae from the Late Paleozoic of Europe. If these two genera are related, this specimen suggests more structural diversity, a wider geographic distribution and a longer geologic range for the Anachoropteridaceae than previously known.

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